The biggest factor in AE’s speed is the clock speed of your processor. Going from 3.8 on the i5 to the 4.2 on the i7 is going to be a noticeable difference.
Some things in After Effects are multithreaded now (like the C4D renderer and the new Camera Shake Deblur) so having multiple cores does help somewhat.
The GPU doesn’t a whole lot for After Effects. However, more and more is being GPU-accelerated. Fractal Noise, Gaussian Blur, Fast Box Blur, Lumetri Color, Sharpen, Brightness and Contrast, Find Edges, Glow, Hue/Saturation, Invert, and Tint have all been given the GPU boost and more has been promised in the future.
Now, some third-party effects render entirely on the GPU or are accelerated heavily by it – Element, Red Giant Universe, Zaxwerks 3d Invigorator, ShapeShifter, etc., for example. So, it really depends on what kind of stuff you will be doing.
Can you get two SSDs in iMacs? AE benefits from having a fast SSD that’s separate from your main drive.
It’s generally considered a good practice to match your RAM sticks. You don’t want to mix sizes.
Also, if you plan to upgrade later, make sure it’s possible first! A lot of Apple products these days have stuff soldered into the main board and can’t be upgraded later, so check before you buy!
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